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A pioneering parasitologist, he spent decades uncovering how human hookworms live, move, and infect their hosts. His work helped shape modern understanding of parasitic disease and earned him wide respect in veterinary and medical research.

by Richard F. Johnston, G. A. Schad
Born in 1928, he became a longtime professor of parasitology at the University of Pennsylvania School of Veterinary Medicine, where he taught and carried out research for more than 35 years.
His best-known work focused on the biology of human hookworms. Colleagues and professional societies remembered him for groundbreaking studies in parasitology, steady support from the National Institutes of Health, and leadership in the field, including service as a past president of the American Society of Parasitologists.
He died in 2009. Beyond the lab, he was also remembered warmly by colleagues, who described a life shaped by curiosity, scholarship, and a deep commitment to science.